Art experts have attributed the enduring intrigue of Leonardo da Vinci's Mona Lisa to everything from having an enigmatic smile, which was never explained, to her association with the rich and powerful families of Europe.
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Ahaving an enigmatic smile, which was never explained
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Bhaving an enigmatic smile, which has never been explained
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Cher enigmatic smile, for which there has never been an explanation
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Dher enigmatic smile, which has never been explained
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Ehaving an enigmatic smile, for which there has never been an explanation
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正确答案: D